英语翻译The bodies of dwarf individuals of mammalianspecies are

英语翻译
The bodies of dwarf individuals of mammalian
species are generally smaller in relation to those of
nondwarf individuals than are the teeth of the dwarf
individuals in relation to those of the nondwarf indi-
viduals.
这个 in relation to 是怎么个用法啊？
还有than 后面为啥跟着个are

我的理解是
哺乳动物中侏儒类动物相对于非侏儒类动物,其体型通常更小,然而其牙齿并不小.
后句话应是than the teeth of the dwarf individuals are in relation to those of the nondwarf indi-
viduals.因为状语从句in relation to those of the nondwarf indi-viduals太长,故用倒装句.
再问： 那倒装之后它为了说什么呢？就是说 这句话里的比较对象 到底是什么？
再答： than的前后分别是bodies和teeth（体型和牙齿）也就是说，相对于非侏儒动物，侏儒动物的体型更小，而牙齿则不小。